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PhD/MPhil Medical Virology / Programme details

Year of entry: 2024

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Programme description

Medical Virology is an important and rapidly expanding field with sensitive molecular techniques leading to the discovery of many new viruses. Dramatic outbreaks of zoonotic virus infections (e.g. Ebola, MERS, SARS, Nipah, etc.,) are regularly seen and changes in climate, society habits, and medical practice have allowed newly emerging and re-emerging viral diseases to spread.

Control of viral disease is challenging; there are currently no antiviral compounds that have truly broad-spectrum activity so preventing and controlling infection is vital. Modern, rapid methods of diagnosis help us to understand the pathogenesis of virus infections, and more and more vaccines and antivirals are being developed and utilized. 

Current research interests include congenital infections, particularly human cytomegalovirus disease, and blood-borne virus infections such as hepatitis C virus, HIV and human polyomaviruses although these areas are not restrictive and a wide range of viral infections are of interest.

During your PhD you will gain practical laboratory experience in molecular virology, cell culture and serology techniques, and learn how these can be applied to both basic science questions and to solve clinical problems.  We work with collaborators within the University, the local hospital Trusts and with a wide ranging European network of clinical virologists.

Our position at the interface between basic research and clinical medicine allows our PhD programme to:

• Enable students to learn how to conduct world-leading research in infection and immunity
• Prepare our students to become independent researchers
• Foster interdisciplinary research

Special features

Training and development

All of our postgraduate researchers attend the Doctoral Academy Training Programme delivered by the Researcher Development team . The programme provides key transferable skills and equips our postgraduate researchers with the tools to progress beyond their research degree into influential positions within academia, industry and consultancy. The emphasis is on enhancing skills critical to developing early-stage researchers and professionals, whether they relate to effective communication, disseminating research findings and project management skills.

Teaching and learning

Applicants are specifically matched with a Primary Supervisor and individual project based on their research interests and background.

International applicants interested in this research area can also consider our PhD programme with integrated teaching certificate .

This unique programme will enable you to gain a Postgraduate Certificate in Teaching and Learning, whilst also carrying out independent research on your chosen project.

Scholarships and bursaries

Funded programmes and projects are promoted throughout the year. Funding is available through UK Research Councils, charities and industry. We also have other internal awards and scholarships for the most outstanding applicants from within the UK and overseas.  

For more information on available the types of funding we have available, please visit the  funded programmes  and  funding opportunities  pages.

What our students say

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• Department of Infectious Disease
• Faculty of Medicine
• Departments

Understanding viruses in the laboratory to tackle them in the clinic

25 group leaders sit within the Section of Virology, most of whom have their laboratory and/or clinical base at the St Mary’s Campus. Their research areas stretch from understanding the molecular details of how viruses manipulate the host cell machinery and subvert immune responses, to optimizing strategies to deliver novel therapies to patients.

The viruses studied within the Section range from the small RNA viruses that cause respiratory infection to the large complex DNA viruses that lead to persistent infections and cancer. There is a strong focus on retrovirology, including strategies for HIV cure, and the early detection and treatment of diseases caused by the human T-lymphotropic virus (HTLV) -1.  Another major focus is to understand the evolution of the influenza virus to improve therapies and vaccines for seasonal influenza and to better predict pandemic emergence. The section has a strong track record in the response to the SARS-CoV-2 pandemic from the development of diagnostics, through determining the impact of viral evolution on the host response to clinical trials. Clinical research spans diagnosis, natural history and pathogenesis studies through early phase clinical trials to international interventions across a range of infections.

To hear more from the virology group follow the @ImperialInfect account .

Research groups and centre leadership

Research groups.

• Interactions between influenza virus and different host species
• Monitoring changes in SARS-CoV-2 and the host response
• Mechanisms and consequence of HTLV-1 integration
• Pathogenesis, diagnosis and treatment of HTLV-associated diseases
• Herpes virus manipulation of the infected cell
• Epstein Barr virus oncogenesis
• Innate responses and therapeutic strategies against Hepatitis viruses
• The Circadian Clock and its impact of virus infections
• Molecular and Cellular Biophysics
• Endogenous retroviruses
• Genetics of infection
• Clinical trials in HIV infection

Section leadership

Professor graham taylor, section head, virology.

Professor Graham Taylor leads the HTLV and the HIV in pregnancy research groups, heads HTLV diagnostics within the Molecular Diagnostic Unit and is a Consultant in the National Centre for Human Retrovirology , the national service for persons infected with HTLVs.

Further information on Professor Taylor

Research leads

Professor wendy barclay.

Professor Wendy Barclay Action Medical Research Chair in Virology

Dr Marta Boffito

Dr Marta Boffito Professor of Practice

Professor Peter Cherepanov

Professor Peter Cherepanov Professor of Molecular Virology

Professor Graham Cooke

Professor Graham Cooke Professor of Infectious Diseases

Dr Rachel Edgar

Dr Rachel Edgar Research Fellow

Dr Charlotte Eve-Short

Dr Charlotte Eve-Short NIHR Academic Clinical Lecturer

Professor Paul Farrell

Professor Paul Farrell Professor of Tumour Virology

Professor Sarah Fidler

Professor Sarah Fidler Professor of HIV and Communicable Diseases

Dr Malick Gibani

Dr Malick Gibani NIHR Academic Clinical Lecturer

Professor Georgios Kassiotis

Professor Georgios Kassiotis Professor of Retrovirology

Dr Goedele Maertens

Dr Goedele Maertens Reader in Virology

Dr Laura Martin-Sancho

Dr Laura Martin-Sancho Lecturer in Molecular Virology

Dr Michael McGarvey

Dr Michael McGarvey Emeritus Reader in Molecular Virology

Professor Peter O'Hare

Professor Peter O'Hare Chair in Virology

Professor Paivi Ojala

Professor Paivi Ojala Visiting Professor

Dr Aileen Rowan

Dr Aileen Rowan Lecturer in Molecular Virology

Professor David Rueda

Professor David Rueda Chair of Molecular and Cellular Medicine

Dr Vanessa Sancho-Shimizu

Dr Vanessa Sancho-Shimizu Reader/UKRI Future Leaders Fellow

Professor Mike Skinner

Professor Mike Skinner Emeritus Professor in Virology

Professor Jonathan Stoye

Professor Jonathan Stoye Professor of Endogenous Retroviruses

Professor Graham Taylor Professor of Human Retrovirology

Dr Lucy Thorne

Dr Lucy Thorne Lecturer in Molecular Virology

Professor Michael Way

Professor Michael Way Professor of Virology

Professor Jonathan Weber

Professor Jonathan Weber Jefferiss Professor of Communicable Diseases and GU Medicine

Dr Rob White

Dr Rob White Senior Lecturer

Professor Alan Winston

Professor Alan Winston Professor of Genito-Urinary Medicine

Professor Xiao-Ning Xu

Professor Xiao-Ning Xu Chair in Human Immunology

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• Research Themes

Immunology & infectious disease

The programme.

The strength and diversity of Oxford's research in immunology and infectious disease means that many projects cut across conventional disciplines, to include aspects of structural biology, protein biology, molecular & cell biology and genomics. Admission is competitive and decided on excellence.

Currently, about half of our students are from outside the UK.

NDM Doctoral Prize Studentships are our fully funded 4 year scholarships open to outstanding students of any nationality, which are advertised each autumn for students who want to start the following October. The current application deadline is early January each year and offers made early February.

Other sources of full funding in this theme include, the Wellcome Trust 4 year PhD programme in Immunology and Infectious Disease , which includes a lab rotation in the first year. Partnership programmes including the NIH-Oxford Scholars Program , Scripps-Oxford and the A*STAR Research Institute in Singapore and various sources of funding for clinicians. The MRC Human Immunology Unit also runs a Doctoral Studentship Competition. A single application covers all funding sources.

Doctoral Projects

• lymphoid development
• innate and adaptive immunity to pathogens and vaccines
• mucosal immunity
• hypersensitivity and autoimmunity
• pathogenic organisms
• vaccinology

Projects relating to Immunology and infectious disease

Training and Career Development

NDM students are part of larger group studying Immunology and Infectious Diseases across Medical Sciences Division's Graduate School. The Oxford Immunology Group organises campus-wide events. Lectures and Seminars cut across departments and collaborations are widespread within the University.

• How to apply
• DPhil in Clinical Medicine
• About the DPhil
• DPhil project themes
• NDM Prize Studentships
• NDM Tropical Network Fund
• CSC-NDM Studentships
• Other funded programmes
• Direct applications
• What to expect
• Student profiles
• For current students
• Past NDM Intern Profiles
• 2024 summer projects
• MSc in Modelling for Global Health
• MSc in International Health & Tropical Medicine
• MSc in Genomic Medicine
• PGDip in Global Health Research

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• Postgraduate study

Postgraduate research opportunities A-Z

• Staff research interests search

Postgraduate research  

Virology PhD/iPhD/MD/MSc (Research)

Virology research is carried out in the MRC-University of Glasgow Centre for Virus Research. Our expertise ranges from molecular virology to in vivo pathogenesis, virus–cell interaction, viral immunology, viral ecology, clinical virology, virus epidemiology, mathematical modelling and bioinformatics.

Research projects

Iphd self-funded projects (november-april).

Our  Integrated PhD combines an MSc and PhD project in a 1+3+1 format. You can select from the below projects and indentify your chosen MSc from the options listed on the project.

Please note that you can apply for the below PhD projects outwith the IPhD route.

Barriers to influenza virus cross-species transmission

Supervisor :  Massimo Palmarini

MSc choice :  Infection Biology (with specialisms) [MSc]

Project outline : Influenza A viruses (IAV) are the cause of a major global health burden in humans and animals. Wild aquatic birds are the main natural reservoir of IAV and are a source of infection for domestic birds, and other species. Spillover of avian IAV into humans results in severe or even lethal disease. These spillover events are typically not followed by extensive human-to-human transmission chains, but they are a risk to global health as they could enable the first step towards human adaptation and the generation of pandemic IAV strains. Multiple barriers have been identified that hamper avian IAV transmission and adaptation in humans including an interferon stimulated gene, BTN3A3, that we recently identified (doi: 10.1038/s41586-023-06261-8). However, several gaps remain in our understanding of how certain avian IAV subtypes/lineages are able to cross the species-barrier and spill over in humans or other mammals. Our laboratory aims to investigate the mechanisms that allow certain avian viruses to spillover into humans using a mixture of virological and evolutionary approaches. Understanding what makes emerging viruses succeed or fail to spillover, and possibly thrive, in human populations are crucial to understand how we manage viral emergence.

Cryogenic correlated light and electron tomography of virally infected cells

Supervisor :  Stephen Carter

Outline and techniques used : We use cryo-CLEM/cryo-electron tomography (cryo-ET) to study the complex relationships between viruses and the host cell during infection.

Our lab has access to the Scottish Centre for Macromolecular Imagining (SCMI) where we use cutting-edge instrumentation, such as the JEOL CRYO ARM 300 electron microscope and the Leica THUNDER Imager EM cryo-CLEM microscope. This technology allows us to target events that happen deep in the cell so we can see more of the context of virions, including their interactions with cellular organelles.

Ultimately, we want to capture the entire virus life cycle, from assembly of its pieces to maturation (with dramatic internal structural changes), to budding, to fusion with a target cell, and then through more transformations as the viral genome passages into the cell’s cytoplasm, all while hijacking the host cell machinery. We work with a range of viruses, including Rift Valley Fever Virus, Bunyamwera virus (BUNV) and herpes simplex virus (HSV).

Project aims:  Developing in situ imaging techniques to image virus infection at high-resolution using cryo-ET. The creation of viral variants that contain and/or can create unnatural amino acids in their own viral proteins and imaging BUNV replication factories using cryogenic focused ion beam (FIB) milling.

• Haney et al. Coinfection by influenza A virus and respiratory syncytial virus produces hybrid virus particles. Nature Microbiology, 2022.
• Carter, S.D., et al. Sci Adv, 2020. 6(14): p. eaay9572.

Understanding virus-virus interactions: from cells to populations

Supervisor :  Pablo Murcia

Background and aims : Respiratory viral infections, including seasonal epidemics and pandemics, cause a major disease burden. Multiple viruses can cause respiratory infections, including influenza viruses, coronaviruses, respiratory syncytial virus, rhinoviruses, human metapneumovirus and parainfluenza viruses, to name but a few. Historically, respiratory viruses have been studied in isolation using a one-virus–one-disease approach. Our laboratory carries out a broad research programme that studies the biology of respiratory viruses using a multi-virus and multi-scale approach (i.e. from cells to populations). In published studies, we combined epidemiological and modelling approaches to reveal the existence of positive and negative interactions between respiratory viruses at the epidemiological scale (1). Using experimental approaches, we showed that interferon responses mediate negative interactions in the human respiratory tract (2, 3). At the cellular level, we were the first to show that virus coinfections can generate infectious hybrid particles (4). Our overarching aim is to determine the processes that drive interactions among respiratory viruses at the population, within-host, and cellular levels.

Techniques to be used : Our group offers a truly multidisciplinary research environment. PhD projects align with the group's overarching research aim and are designed around the students' training needs. Wet lab projects include cell culture, classical virology, immunostaining, microscopy, and imaging (2-4) as well as serological assays (5, 6). Dry projects combine epidemiology (7, 8), evolutionary biology, bioinformatics (9) and modelling (1, 10).

• S. Nickbakhsh et al., Virus-virus interactions impact the population dynamics of influenza and the common cold. Proc Natl Acad Sci U S A 116, 27142-27150 (2019).
• K. Dee et al., Human Rhinovirus Infection Blocks Severe Acute Respiratory Syndrome Coronavirus 2 Replication Within the Respiratory Epithelium: Implications for COVID-19 Epidemiology. J Infect Dis 224, 31-38 (2021).
• K. Dee et al., Influenza A and Respiratory Syncytial Virus Trigger a Cellular Response That Blocks Severe Acute Respiratory Syndrome Virus 2 Infection in the Respiratory Tract. J Infect Dis 227, 1396-1406 (2023).
• J. Haney et al., Coinfection by influenza A virus and respiratory syncytial virus produces hybrid virus particles. Nat Microbiol 7, 1879-1890 (2022).
• E. C. Hughes et al., Severe Acute Respiratory Syndrome Coronavirus 2 Serosurveillance in a Patient Population Reveals Differences in Virus Exposure and Antibody-Mediated Immunity According to Host Demography and Healthcare Setting. J Infect Dis 223, 971-980 (2021).
• M. Manali et al., SARS-CoV-2 Evolution and Patient Immunological History Shape the Breadth and Potency of Antibody-Mediated Immunity. J Infect Dis 227, 40-49 (2022).
• S. Nickbakhsh et al., Extensive multiplex PCR diagnostics reveal new insights into the epidemiology of viral respiratory infections. Epidemiol Infect 144, 2064-2076 (2016).
• S. Nickbakhsh et al., Epidemiology of Seasonal Coronaviruses: Establishing the Context for the Emergence of Coronavirus Disease 2019. J Infect Dis 222, 17-25 (2020).
• F. Thorburn et al., The use of next generation sequencing in the diagnosis and typing of respiratory infections. J Clin Virol 69, 96-100 (2015).
• C. Mair et al., Estimation of temporal covariances in pathogen dynamics using Bayesian multivariate autoregressive models. PLoS Comput Biol 15, e1007492 (2019).

The MRC-University of Glasgow Centre for Virus Research (CVR) sits within the School of Infection and Immunity . The CVR is the largest virology-focussed research centre in the UK and brings together a critical mass of researchers studying human and animal viruses and viral diseases.

The CVR provides excellent facilities and opportunities for cross-disciplinary projects and the delivery of a comprehensive programme of training in contemporary, multi-disciplinary, virology research. The Centre includes research programmes in arboviruses, Epstein Barr virus, feline calicivirus, herpes viruses, hepatitis C virus, influenza, retroviruses and papillomaviruses.

 Cross cutting research themes and expertise include:

• antiviral immunity
• virus discovery
• viral bioinformatics, mathematical modelling and genomics to guide new approaches to the understanding and management of viral infections
• structural biology/cryo-electron microscopy and viral evolutionary dynamics
• molecular virology to in vivo pathogenesis
• virus-cell interactions
• viral immunology
• viral ecology
• viral oncology
• clinical and veterinary virology
• viral diagnostics 
• virus epidemiology

Our excellent facilities underpin a bench to bedside approach that will equip you with training complementary to a range of career options, and you can tailor your study pathway to the precise aspects of infection and immunology that suit your objectives. Through their research interests in drug development, vaccines and diagnostics, many of our project supervisors have strong links with industry.

Study options

• Duration:  3/4 years full-time; 5 years part-time

Individual research projects are tailored around the expertise of principal investigators.

Integrated PhD programmes (5 years)

Our  Integrated PhD  allows you to combine masters level teaching with your chosen research direction in a 1+3+1 format. 

International students with MSc and PhD scholarships/funding do not have to apply for 2 visas or exit and re-enter the country between programmes. International and UK/EU students may apply.

Taught masters level modules are taken alongside students on our masters programmes. Our research-led teaching supports you to fine tune your research ideas and discuss these with potential PhD supervisors. You will gain a valuable introduction to academic topics, research methods, laboratory skills and the critical evaluation of research data. Your grades must meet our requirements in order to gain entry on to your pre-selected PhD research project. If not, you will have the options to pay outstanding MSc fees and complete with masters degree only.

Years 2, 3 and 4

PhD programme with research/lab work, completing an examinable piece of independent research in year 4.

Thesis write up.

MSc (Research)

• Duration : 1 year full-time; 2 years part-time

MD (Doctor of Medicine)

• Duration : 2 years full-time; 4 years part-time (for medically-qualified graduates only)

Entry requirements

A 2.1 Honours degree or equivalent.

English language requirements

For applicants whose first language is not English, the University sets a minimum English Language proficiency level.

International English Language Testing System (IELTS) Academic module (not General Training)

• 6.5 with no subtests under 6.0
• Tests must have been taken within 2 years 5 months of start date. Applicants must meet the overall and subtest requirements using a single test.

Common equivalent English language qualifications accepted for entry to this programme:

Toefl (ibt, my best or athome).

• 79; with Reading 13; Listening 12; Speaking 18;Writing 21
• Tests must have been taken within 2 years 5 months of start date. Applicants must meet the overall and subtest requirements , this includes TOEFL mybest.

Pearsons PTE Academic

• 59 with minimum 59 in all subtests

Cambridge Proficiency in English (CPE) and Cambridge Advanced English (CAE)

• 176 overall, no subtest less than 169

Oxford English Test

• Oxford ELLT 7
• R&L: OIDI level no less than 6 with Reading: 21-24 Listening: 15-17
• W&S: OIDI level no less than 6

Trinity College Tests

Integrated Skills in English II & III & IV: ISEII Distinction with Distinction in all sub-tests.

University of Glasgow Pre-sessional courses

Tests are accepted for 2 years following date of successful completion.

Alternatives to English Language qualification

• students must have studied for a minimum of 2 years at Undergraduate level, or 9 months at Master's level, and must have complete their degree in that majority-English speaking country  and  within the last 6 years
• students must have completed their final two years study in that majority-English speaking country  and  within the last 6 years

For international students, the Home Office has confirmed that the University can choose to use these tests to make its own assessment of English language ability for visa applications to degree level programmes. The University is also able to accept UKVI approved Secure English Language Tests (SELT) but we do not require a specific UKVI SELT for degree level programmes. We therefore still accept any of the English tests listed for admission to this programme.

Pre-sessional courses

The University of Glasgow accepts evidence of the required language level from the English for Academic Study Unit Pre-sessional courses. We also consider other BALEAP accredited pre-sessional courses:

• School of Modern Languages and Cultures: English for Academic Study
• BALEAP guide to accredited courses

Fees and funding

• UK: £4,786
• International & EU: £30,240

Prices are based on the annual fee for full-time study. Fees for part-time study are half the full-time fee.

Irish nationals who are living in the Common Travel Area of the UK, EU nationals with settled or pre-settled status, and Internationals with Indefinite Leave to remain status can also qualify for home fee status.

• Fee status and policies

Alumni discount

We offer a 20% discount to our alumni on all Postgraduate Research and full Postgraduate Taught Masters programmes. This includes University of Glasgow graduates and those who have completed Junior Year Abroad, Exchange programme or International Summer School with us. The discount is applied at registration for students who are not in receipt of another discount or scholarship funded by the University. No additional application is required.

Possible additional fees

• Re-submission by a research student £540
• Submission for a higher degree by published work £1,355
• Submission of thesis after deadline lapsed £350
• Submission by staff in receipt of staff scholarship £790

Depending on the nature of the research project, some students will be expected to pay a bench fee (also known as research support costs) to cover additional costs. The exact amount will be provided in the offer letter.

The iPhD  is not supported by University of Glasgow Scholarship/Funding

• BBSRC Doctoral Training Partnerships
• External funding information

The College of Medical, Veterinary and Life Sciences Graduate School provides a vibrant, supportive and stimulating environment for all our postgraduate students. We aim to provide excellent support for our postgraduates through dedicated postgraduate convenors, highly trained supervisors and pastoral support for each student.   Our overarching aim is to provide a research training environment that includes:

• provision of excellent facilities and cutting edge techniques
• training in essential research and generic skills
• excellence in supervision and mentoring
• interactive discussion groups and seminars
• an atmosphere that fosters critical cultural policy and research analysis
• synergy between research groups and areas
• extensive multidisciplinary and collaborative research
• extensive external collaborations both within and beyond the UK 
• a robust generic skills programme including opportunities in social and commercial training

Research environment

If you study with us, you will join a large community of postgraduate taught and research students. Our School brings together basic, applied, clinical and translational researchers to study infection with a focus on the viral, parasitic and bacterial pathogens of both humans and animals, and immunology and inflammation with a focus on chronic inflammatory diseases.

Despite the continual development of new therapies, antibiotics and vaccines, chronic inflammatory and infectious diseases still pose persistent health threats. We aim to:

• understand the basic science of the immune systems and how the immune system can inturn affect disease outcome understand the biology of parasites, viruse and bacteria and the interactions with their hosts, that in turn leads to high levels of infectious diseases worldwide
• develop therapies (drugs and vaccines) targeted on these processes
• explore new treatments and strategies in clinical and translational medicine

Research centres

• MRC-University of Glasgow Centre for Virus Research
• Wellcome Centre for Integrative Parasitology
• ARUK Centre for Experimental Arthritis Research

We offer a wide range of cutting-edge research facilities, including core facilities in fluorescence activated cell sorting analysis, histology and state-of-the-art imaging. In addition, we offer the IVIS imaging system, high content screening microscopy, mass spectrometry, an X-ray capable FX Pro bioluminescence imaging system and a protein purification service. Also available are a wide range of molecular, immunological and biochemical analysis tools.

How to apply

Identify potential supervisors.

All Postgraduate Research Students are allocated a supervisor who will act as the main source of academic support and research mentoring. You may want to identify a potential supervisor and contact them to discuss your research proposal before you apply. Please note, even if you have spoken to an academic staff member about your proposal you still need to submit an online application form.

You can find relevant academic staff members with our staff research interests search .

IPhD applicants do not need to contact a supervisor, as you will choose from a list of IPhD projects. Each project has named supervisors.

Gather your documents

Before applying please make sure you gather the following supporting documentation:

• Final or current degree transcripts including grades (and an official translation, if needed) – scanned copy in colour of the original document.
• Degree certificates (and an official translation, if needed): scanned copy in colour of the original document
• Two references on headed paper and signed by the referee. One must be academic, the other can be academic or professional [except iPhD applicants, where only one academic or professional reference is required]. References may be uploaded   as part of the application form or you may enter your referees contact details on the application form. We will then email your referee and notify you when we receive the reference.  We can also accept confidential references direct to  [email protected] , from the referee’s university or business email account.
• Research proposal, CV, samples of written work as per requirements for each subject area. iPhD applicants do not need to submit any of these as you will start your programme by choosing a masters.
• Completed  College of MVLS Postgraduate Research Cover Letter

Notes for iPhD applicants

• add 'I wish to study the MSc in (select MSc from IPhD project choices) as the masters taught component of the IPhD' in the research proposal box
• For supervisor name, please ensure you write the named supervisors from your chosen IPhD project.

Before you apply

PhD/MSc/MD: email  [email protected]

iPhD: email  [email protected]

After you have submitted your application

PhD/MSc/MD/iPhD:  contact our Admissions team

Any  references  may be submitted by email to:  [email protected]

Epidemiology and Public Health MPhil/PhD

London, Bloomsbury

Our PhD programme aims to equip the next generation of experts with the necessary tools to address major 21st-century health challenges and deliver real-world impact.

UK tuition fees (2024/25)

Overseas tuition fees (2024/25), programme starts, applications accepted.

Prospective students should apply at least four months before their intended start date. If you require a visa we recommend allowing for more time.

• Entry requirements

A minimum of an upper second-class UK Bachelor’s degree and/or a Master’s degree (preferably with a merit or distinction) in a relevant discipline, or an overseas qualification of an equivalent standard.

The English language level for this programme is: Level 2

UCL Pre-Master's and Pre-sessional English courses are for international students who are aiming to study for a postgraduate degree at UCL. The courses will develop your academic English and academic skills required to succeed at postgraduate level.

Further information can be found on our English language requirements page.

Equivalent qualifications

Country-specific information, including details of when UCL representatives are visiting your part of the world, can be obtained from the International Students website .

International applicants can find out the equivalent qualification for their country by selecting from the list below. Please note that the equivalency will correspond to the broad UK degree classification stated on this page (e.g. upper second-class). Where a specific overall percentage is required in the UK qualification, the international equivalency will be higher than that stated below. Please contact Graduate Admissions should you require further advice.

About this degree

The Epidemiology and Public Health research degree programme is based within UCL's Institute of Epidemiology and Health Care.

You will join an extremely engaging PhD programme, which currently supports a vibrant community of over 100 students from all over the world.

This programme offers you the opportunity to join a multidisciplinary research department with experts who focus on a wide range of public health concerns such as:

• Infectious disease monitoring, prevention and control
• Social determinants of health 
• Dental public health
• Disability across the globe
• Health and life expectancy in socially excluded groups
• Health surveys and longitudinal cohort studies

You will be supported to gain a wealth of skills, experience and networking opportunities that are relevant for a wide range of future careers, both within academia and more widely in the healthcare, industry and governmental sector.

You will receive direct supervision from world-leading academics based on UCL's campus in the heart of London. All PhD students are supported by supervisory panels of multidisciplinary staff, as well as student mentors, and departmental graduate tutors.

Who this course is for

This programme provides training for those looking for a career in epidemiology, public health and health care policy, either in academia, industry or public health practice. It is offered full time or part time, the latter option being suitable for those who continue in employment while gaining a research qualification.

What this course will give you

This programme will provide you with access to a wide range of supporting opportunities that will increase your ability to develop transferable skills, that are sought after by national and international employers.

Skills Development programme

UCL's DocSkills Development Programme is open to all PhD students at UCL and it offers an extensive list of development opportunities. The purpose of the programme is to give you the opportunity to expand your research and transferable skills in order to support your research, professional development and employability. Find out more about UCL's DocSkills Development programme .

Mentoring programme

We offer a Peer-Level Research Student Mentoring Scheme and provide all new research degree students with a peer-mentor upon arrival. The mentor you are allocated will generally be a 2nd or 3rd year PhD student.

We take career support very seriously at UCL and you can find out more within the programmes Careers and Employability section .

Early Career Researchers Forum

The Institutes’s Early Career Researcher Forum (ECF) is an ongoing and expanding programme of regular seminars for postdoctoral researchers and PhD students. Previous events focused on how to apply for jobs in academia, authorship and publishing, patient and public involvement in research, using social media in research (including film), how PhD students can best prepare for their viva, and oral and poster presentations.

Journal Club

A PhD and junior researcher journal club group which meets in an informal setting. With the aim of improving student's critical analysis and methodology skills, the Journal Club discusses papers from a broad range of topics on health promotion and disease prevention. The club is run by students and facilitated by a senior member of academic staff within the Institute.

Lunchtime Seminar Series

The Lunchtime Seminar series is delivered by the department's research groups and PhD students. The seminars typically take place once a month.

PhD Poster Competition

Every year we run a PhD student poster competition which is an opportunity to share students work with members of staff, outsiders who will be visiting for our Open Day and fellow students.

3-minute thesis competition (3MT)

The Institute runs an annual 3-minute thesis competition which is an academic competition that challenges PhD students to describe their research within three minutes to a general audience. 3MT celebrates the discoveries made by research students and encourages them to communicate the importance of their research to the broader community. It's a great opportunity for students to practice their presentation skills, meet other candidates and have a chance to win prize money.

Teaching Opportunities for PhD Students

The Institute is committed and recognises the importance of providing postgraduate teaching assistant opportunities for PhD students, so they can gain valuable experience during their studies at UCL. Formal is provided and a wide range of teaching opportunities are offered each term.

The foundation of your career

We produce graduates with the skills and knowledge sought after by government departments and public sector organisations worldwide, as well as leading academic institutions.

Employability

This research degree programme aims to provide excellent and challenging training for exceptional students, so that they may successfully pursue careers in:

• Local and central government
• Public health organisations
• Hospitals and clinical trial units
• Academic researchers and university lecturers
• Policy management
• Government advisors
• Charity and non-governmental sectors

This degree can be varied and may have an international dimension, including fieldwork carried out abroad, setting up a study within the UK or other countries, or using secondary data from some of the UK’s largest, most comprehensive longitudinal datasets many of which are housed within the institute.

You will also have access to UCL's Doctoral Skills Development programme, which will enable you to expand your research and transferable skills in order to support your research, professional development and employability.

The Institute’s research departments collaborate with third sector and governmental organisations, as well as members of the media, both nationally and internationally to ensure the highest possible impact of their work beyond the academic community. Students are encouraged to take up internships with relevant organisations where funding permits. Members of staff also collaborate closely with academics from leading institutions globally.

Teaching and learning

Learning is mostly self-directed with input from PhD supervisors. The training and development programme for each student is overseen and supported by a Thesis Committee panel, appointed by the supervisory team.

PhD final assessment is by means of a thesis, which should demonstrate your ability to pursue original research based upon a good understanding of the research techniques and concepts appropriate to the discipline.

Your thesis must also represent your distinct and significant contribution to the subject, either through the discovery of new knowledge, through the connection of previously unrelated facts, the development of a new theory, or the revision of older views.

Your thesis should reflect the exercise of critical judgement with regard to both your own work and that of other scholars in the field.

You are required to register initially for the MPhil degree with the expectation of transfer to PhD after successful completion of an upgrade viva. For a successful upgrade to PhD, you prepare a written report, give an oral presentation and pass an oral examination.

For a PhD award, you will be examined on your submitted thesis, this involves a viva (an oral exam) with two independent examiners.

Contact hours and hours of self-study are agreed between the student and the supervisor at the beginning of their research degree and should be reviewed on a regular basis. Full-time postgraduate research students are expected to work a minimum of 36.5 hours per week on their project. With agreement of their supervisors, contact time can be on-site or remote working depending upon the nature and stage of the project. PGR students can have the opportunity to access UCL facilities ‘out of hours’ including weekends and holidays during their period of registration. Students will have research meetings with their supervisors at least once per month. Full-time Research students can take 27 days of annual leave, plus eight days of Bank holidays and six UCL closure days.

Research areas and structure

Our research focuses on a wide range of public health concerns such as heart disease; dental public health; mental health and well-being; and child development and ageing. Our research has significant real-world impact, informing policy both in the United Kingdom and around the world, and the wider public understanding of health inequalities.

Find out more.

Research environment

UCL is among the world's top ten universities (QS World University Rankings 2024) with a reputation for high-quality research. Located in the heart of London, it is a stimulating and exciting environment in which to study.

• UCL is rated No.1 for research power and impact in medicine, health, and life sciences (REF 2021)
• UCL is ranked 6th in the world for public health (ShanghaiRanking's 2023 Global Ranking of Academic Subjects)
• UCL is ranked 9th in the world as a university (QS World Rankings 2024)

UCL can bring the full power of a multi-faculty university to bear on discussions of population health, involving academics from the wide range of disciplines necessary to tackle some of the most difficult issues in public health.

More specifically, the UCL's Institute of Epidemiology and Health Care brings together four research departments, whose interests span the life course from childhood to old age, and research from the origins of disease to the development of innovative interventions.

The four research Departments are:

• Applied Health Research
• Behavioural Science and Health
• Epidemiology and Public Health
• Primary Care and Population Health

Full-time students are normally registered for a minimum of 3 years. 

All students initially register for the MPhil degree before being upgraded (typically early in the 2nd year) to the PhD degree. For a successful upgrade to a PhD, students must prepare a written report, give an oral presentation and pass an oral examination. 

Students are regularly monitored to ensure that they are making good progress and that supervisory arrangements are satisfactory to both the student and supervisor. Once the student has completed their research and submitted their thesis, they have a viva (an oral exam) with two examiners.

Part-time students are required to register for a minimum of 5 years.

Accessibility

Details of the accessibility of UCL buildings can be obtained from AccessAble accessable.co.uk . Further information can also be obtained from the UCL Student Support and Wellbeing team .

Fees and funding

Fees for this course.

The tuition fees shown are for the year indicated above. Fees for subsequent years may increase or otherwise vary. Where the programme is offered on a flexible/modular basis, fees are charged pro-rata to the appropriate full-time Master's fee taken in an academic session. Further information on fee status, fee increases and the fee schedule can be viewed on the UCL Students website: ucl.ac.uk/students/fees .

Additional costs

There may be additional costs related to participating in scientific meetings, conferences, short courses, or data collection/access, but these are all optional.

The students can apply for financial support provided by the Institute of Epidemiology and Health Care to participate in conferences.

For more information on additional costs for prospective students please go to our estimated cost of essential expenditure at Accommodation and living costs .

Funding your studies

View the Institute of Epidemiology and Health Care's Studentship and Funding webpage for opportunities.  

The following studentships have been received in previous years: ESRC, MRC, MRC/ESRC, NIHR, Cancer Research UK, Diabetes UK, Wellcome Trust, European Union and British Heart Foundation.

For a comprehensive list of the funding opportunities available at UCL, including funding relevant to your nationality, please visit the Scholarships and Funding website .

We accept research degree applications throughout the year. Please note: essential information about the application process can be found on the website .

Please note that you may submit applications for a maximum of two graduate programmes (or one application for the Law LLM) in any application cycle.

Choose your programme

Please read the Application Guidance before proceeding with your application.

Year of entry: 2024-2025

Got questions get in touch.

Institute of Epidemiology and Health Care

[email protected]

UCL is regulated by the Office for Students .

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• Funding your Master's

Funded programmes

We offer several specialised PhD training programmes that come with funding.

All of our fully funded programmes are displayed below including our prestigious doctoral training partnerships and industry funded schemes.

Some of these programmes offer interdisciplinary or structured training during the first year or placements outside of the lab.

Alternatively, you can search our live project listings to find advertised fully funded projects or visit our funding opportunities page  to browse the fully funded scholarships and awards we offer.

Excellence in Diversity PhD Scholarships

Fully funded studentships for candidates from groups under-represented in postgraduate research.

More about the Excellence in Diversity scholarships

Joint scholarships with the China Scholarship Council (CSC)

Scholarships for Chinese PhD students through a joint scheme between the University and the CSC.

More about joint scholarships with the CSC

BBSRC DTP studentships

The BBSRC Doctoral Training Partnership provides an opportunity to experience cutting-edge research in the biosciences.

More about BBSRC DTP studentships

MRC DTP studentships

The MRC Doctoral Training Partnership provides an outstanding environment for PhDs in the biomedical sciences.

More about MRC DTP studentships

British Heart Foundation PhD

Our 4-year BHF PhD provides an intensive introduction to cardiovascular research.

More about the British Heart Foundation PhD

Wellcome Trust Immunomatrix in Complex Disease PhD

A 4-year PhD programme at the interface of immunology and matrix biology research.

More about the Wellcome Trust Immunomatrix in Complex Disease PhD

CRUK Manchester Centre PhD Training Scheme

Develop a career in world-leading cancer research with clinical, MB-PhD and non-clinical PhD opportunities funded by the Cancer Research UK Manchester Centre.

More about the CRUK Manchester Centre PhD Training Scheme

CRUK Manchester Centre MB-PhD

Combining medical training with research training in the area of cancer sciences.

More about the CRUK Manchester Centre MB-PhD

CRUK Manchester Institute PhD studentships

The Cancer Research UK Manchester Institute have PhD projects on offer spanning the whole spectrum of cancer research.

More about the CRUK Manchester Institute PhD studentships

ESRC North West Social Science Doctoral Training Partnership

ESRC funding for postgraduate training in the social sciences is available through the North West Social Sciences DTP.

More about the ESRC North West Social Science DTP

Social-Biological CDT

PhD studentships are available in an innovative new centre for doctoral training in biosocial research.

More about the Social-Biological CDT

HDR UK/Turing PhD Programme in Health Data Science

PhD studentships are available for research within health data science.

More about the HDR UK/Turing PhD Programme in Health Data Science

Manchester Biomedical Research Centre (BRC) PhD studentships

Studentships for clinical and non-clinical PhD projects in cancer, inflammation and other areas with the Manchester BRC.

More about the Manchester BRC PhD studentships

President's Doctoral Scholar awards

Studentships for outstanding international (non-UK) candidates seeking PhD opportunities.

More about the President's Doctoral Scholar awards

Dual Award PhD with the University of Melbourne

Apply for a project from the Dual Award PhD programme between Manchester and Melbourne.

More about the Dual Award PhD with the University of Melbourne

Kennedy Trust IMPACT Inflammation MB-PhD

Combining the vocational training of a medical degree with the research expertise of a PhD.

More about the Kennedy Trust IMPACT Inflammation MB-PhD

The Boots Company Collaborative Training Partnership (CTP)

Undertaking world-class bioscience research in skin health.

More about the The Boots Company CTP

Funded projects

Our projects database individually lists the studentship opportunities available.

Search the database

Funding opportunities

Find out about the other sources of financial support available for UK and international applicants.

See funding opportunities

• Postgraduate Research

Vaccinology PhD / MPhil / MD

• Part time available: yes

Studying in:

• institute-of-infection-veterinary-and-ecological-sciences
• Faculty of Health and Life Sciences

We have established a Centre for Global Vaccine Research to integrate world-leading multidisciplinary research and training in laboratory science, clinical trials and epidemiology within the Institute of Infection and Global Health, and reaching across the Faculty of Health and Life Sciences and partners around the world.

Why study with us?

I joined an amazing community of internationally known researchers in the most relevant areas of infectious diseases. During my PhD at the Institute I had many opportunities to develop both my scientific knowledge and skills as well as my personal capacities such as communication and networking. Raquel Medialdea-Carrera, Vaccinology PhD student

years of leading international reputation.

research students.

annual research income.

The University of Liverpool’s Institute of Infection and Global Health was established to bring together leading medical, veterinary and basic science researchers from across the Faculty of Health and Life Sciences. 

The Centre for Global Vaccine Research currently works across Europe, Asia, Africa and more recently South America. Our research spans the breadth of vaccine research from development to implementation and evaluation.

The key aims of this research are to:

• Develop and evaluate novel vaccines against major global causes of illness and death
• Better understand and improve the performance of existing vaccines
• Generate evidence to inform vaccine policy nationally and internationally
• Share pre-clinical expertise amongst human and veterinary vaccinologists
• Train and educate a new generation of vaccine scientists.

Our research interests

We particularly welcome research proposals that match those of our researchers, including:

• Flavivirus Vaccine Development (including Zika virus) 
• Rotavirus Vaccines 
• Pneumococcal Vaccines 
• Universal Influenza Vaccines 
• Onchocerciasis Vaccines  
• Streptococcal vaccines
• Enteric vaccines (including norovirus and enteric bacteria)
• Veterinary vaccines (including Fasciola, Trypanosomiasis and avian respiratory viruses) 
• Herpes viruses

Research themes

The Centre for Global Vaccine Research has six research themes covering key methodological disciplines within vaccinology and areas of strength shared by the Centre partners. These disciplines can be applied to both medical and veterinary vaccines.  The themes are:

• Disease epidemiology & Clinical trials
• Vaccine adjuvants
• Microbial ecology
• Antigen discovery & vaccine development
• Pre-clinical assessment models
• Immune correlates of protection

Within IGH there is access to a wide array of facilities for conducting vaccine research including  immunology, in vivo animal models, tissue culture and microbial propogation (including category 3 pathogens). In the wider university there is the technology directorate which gives access to various imaging and omics platforms. There is a clinical trials unit and phase 1 trial facility in the neighbouring Royal Liverpool hospital. There are established links with local primary care and public health England. The vaccine centre has strong links to overseas sites in Africa, Asia and more recently South America. We have a particularly strong connection to Malawi where many former clinical and non-clinical PhDs have spent time in the Wellcome Trust Major Overseas program Laboratories.

Research groups

Details of research groups undertaking research in vaccinology include:

• Flavivirus Vaccine Development             
• Rotavirus and Enteric Vaccines               
• Pneumococcal Vaccines                       
• Universal Influenza Vaccines              
• Onchocerciasis Vaccines                       
• Streptococcal vaccines                                
• Veterinary vaccines                                    
• Herpesviruses                                               

Study options and fees

The fees stated in the table above exclude potential research support fees also known as ‘bench fees’. You will be notified of any fee which may apply in your offer letter.

* Please note that if you are undertaking a PhD within the Faculty of Science and Engineering the fee you pay, Band A or Band B, will reflect the nature of your research project. Some research projects incur a higher fee than others e.g. if you are required to undertake laboratory work. You will be informed of the fee for your programme in your offer letter.

^ Self-funded, full-time international students studying a PhD programme classified as Band A will receive a £2,000 reduction in their fees for the first year only.

Entry requirements

Students will normally have a minimum of a 2.1 class honours degree in a relevant biological science subject, or an equivalent medical, veterinary or dental qualification. Applicants are selected on the basis of their curriculum vitae, qualifications and referees’ reports, together with their perceived ability to complete the programme successfully.

English language requirements

How to apply.

Research degree applications can be made online.  You'll also need to ensure that you have funding to cover all fees.

Applications are  open all year round .

More about applying for research degrees

Apply online

Before you apply, we recommend that you identify a supervisor and develop a research proposal

Find a supervisor

• Prof Nigel Cunliffe                      
• Prof Ituriza-Gomara
• Prof Neil French
• Prof Aras Kadioglu
• Prof Tom Solomon
• Dr Lance Turtle
• Dr Qibo Zhang
• Dr Robin Flynn
• Dr Andrew Jackson
• Dr Dan Neill

Need help finding a supervisor? Contact us

• IVES PGR Administrator
• Email: [email protected]
• Phone: +44 (0)151 794 8032

Related studentships: self-funded and funded PhD projects

Find a scholarship.

We offer a range of scholarships to help you meet the costs of studying a research degree.

See scholarships

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PhD in Virology: Fellowships, Jobs, Positions

Find a PhD in virology and its related areas in the UK, USA, Europe, Canada, Australia, New Zealand, etc. Virology PhD programs, scholarships, research projects, fellowships, courses, and positions are available in European countries, Germany, Finland, Belgium, Netherlands, Switzerland, Sweden, Norway, Denmark, Italy, etc. Here you can also find studentships, jobs, courses, offers, and fully funded opportunities for international students abroad.

Fellowships for PhD in Virology

Research associate, school of medicine and health sciences (smhs), george washington university, usa, february 21, 2024, the uni jobs.

Published Date: 02/14/2024. Posting Details I. DEPARTMENT INFORMATION Job Description Summary: The laboratory of Dr. David Watkins is looking for a talented Research Associate who is interested in working in the development and efficacy testing of vaccines, diagnostics, and therapeutic drugs against virus and other diseases including cancers. The Watkins’ …

PhD student for 4 years. The research work is biomedical science with a focus on molecular parasitology, Swedish University of Agricultural Sciences, Sweden, Europe

February 5, 2024, ph.d , scholarships.

Last Date: 2024-03-01. Department of Animal Biosciences. (HBIO) HBIO is part of the Faculty of Veterinary Medicine and Animal Sciences and is responsible for education, research and postgraduate studies in several basic and applied scientific areas. These areas include anatomy, physiology, biochemistry, pathology, pharmacology, toxicology, breeding and genetics, immunology, bacteriology, …

Students Wanted for Exciting Summer Jobs at the School of Chemical Engineering, Aalto University, Finland, Europe

December 11, 2023.

Last Date: 18.2.2024. The open positions are in all the departments of our School: Department of Bioproducts and Biosystems (Bio2), the Department of Chemical and Metallurgical Engineering and the Department of Chemistry and Materials Science. How to apply? Please submit your application through our recruitment system by using the “Apply” link on this …

Research Assistant / Associate in Virus Enzyme Biochemistry (Fixed Term), University of Cambridge, Cambridge, UK

September 7, 2023.

Last Date: 30 October 2023. We have an exciting opportunity for a postdoctoral research scientist* to study the physical properties of viral enzymes that allow them to evolve resistance to anti-viral drugs. This project will use advanced computation biology approaches, enzymology and structural biology to define structurally invariant regions of …

All about All Sciences - Sign up for Updates

Studentships and doctoral training

Get a studentship to fund your doctorate.

UKRI studentships offer funding for doctoral research. They also offer you access to training, networking and development opportunities to help you build a research and innovation career.

Our expectations for research organisations, supervisors and students are set out in the statement of expectations for doctoral training .

You could get:

• a minimum stipend of £19,237 per year for your living costs, which is paid to you in regular instalments
• support for your tuition fees (minimum £4,786 per year)

The stipend is usually non-taxable and does not need to be paid back. Some research organisations may offer more if you study in London, or they or one of their collaborators might decide to top up the payment. This will be outlined in the studentship advert from the research organisation.

We normally pay the support for tuition fees directly to your research organisation.

The levels given here are for the academic year 2024 to 2025. UKRI’s approach to doctoral stipend and fee levels will be reviewed through the  new deal for postgraduate research .

Additional support for your doctoral studies

As a UKRI-funded doctoral student, you may be able to access additional funding to cover the cost of other related training and development opportunities.

This could include:

• conference attendance
• language training
• overseas research visits
• internships or placements with a non-academic partner

The availability of support will depend on the research organisation and the training grants they have on offer. You should contact the research organisation you are interested in applying to, to find out what you could get.

Extra support if you have a disability

If you have a disability, you may be entitled to a Disabled Students’ Allowance (DSA) on top of your studentship.

You should speak to your research organisation’s disability advisor to assess your needs. They can help put the right support in place, including a DSA application if necessary. You cannot claim DSA directly from UKRI.

DSA helps to cover the cost of any additional support that a person studying for a doctorate might need as a result of a disability, mental health problem or specific learning difficulty.

The allowance covers:

• non-medical personal assistance
• specialist equipment
• extra travel costs
• general expenses

Find out more about DSA in our framework .

If you are a research organisation you can download claim forms and guidance for DSA .

Who can apply

Any prospective doctoral student wishing to study at a UK research organisation, including prospective international students, can apply for a UKRI studentship.

All UKRI-funded doctoral students will be eligible for the full award, both the stipend to support living costs, and home-level fees at the UK research organisation rate.

How to find opportunities

Many UK research organisations offer some form of studentship funding from UKRI. These opportunities will depend on the subject you want to study and will normally be advertised by the research organisations.

Research organisations may have additional opportunities that do not involve UKRI. UKRI supports around 20% of all UK-based postgraduate researchers. You should speak to the research organisation you are interested in to find out what studentships are available.

You could also consider using a specialist website like   FindaPhD  to look for opportunities.

When to apply

Research organisations set their own deadlines for applications.

Many open for applications early in the academic year and close in January or February. This is not a hard and fast rule. It is important that you check the deadlines for the research organisation where you want to study.

How to apply

You cannot apply to UKRI for a studentship. You must contact the research organisation you are interested in studying with and use their application process.

For doctoral students who are already studying with a studentship, there are opportunities to get additional funding to support placements that are separate from your doctorate.  Find training and development opportunities .

Last updated: 14 February 2024

This is the website for UKRI: our seven research councils, Research England and Innovate UK. Let us know if you have feedback or would like to help improve our online products and services .

65 phd-virology positions in europe

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PhD /Masters Scholarship, WENS, KIT (Fall 2024)

), Unmanned Mobility (UAV, UGV, USV, UUV), etc. For Fall 2024, PhD and MS- PhD Integrated will be prioritized. The scholarship includes full tuition fee until graduation, medical insurance and a monthly stipend

PhD Candidate in the Cultures of Palestine Solidarity in Europe

related to staff position within a Research Infrastructure? No Offer Description You are invited to apply for a PhD position on the cultural histories of Palestine solidarity and cultural activism in Europe

17 fully funded PhD positions in Chemistry

funded by an EU programme Is the Job related to staff position within a Research Infrastructure? No Offer Description The Vienna Doctoral School in Chemistry offers 17 fully funded PhD positions to be

PhD Vacancy - A transdisciplinary multi-scalar approach to foster mental health in Europe by means of nature-based therapies

The PhD project will be embedded within the HEurope GreenME project (September 2023-August 2027). All supervisors are involved in the project, which is coordinated by H. Cole and M. Triguero-Mas. GreenME

PhD Position: Chinese Migrants' Selves in Europe

Job related to staff position within a Research Infrastructure? No Offer Description Are you looking for a challenging position in a dynamic setting? Research school ARTES currently has a vacant PhD

position within a Research Infrastructure? No Offer Description The Vienna Doctoral School in Chemistry offers 17 fully funded PhD positions to be filled between early summer and the end of 2024

Are you looking for a challenging position in a dynamic setting? Research school ARTES currently has a vacant PhD position as part of the project “Me, Myself, and I – Chinese Migrants’ Selves in

PhD position in the EU Horizon Europe project REDUCE

Programme? HE Is the Job related to staff position within a Research Infrastructure? No Offer Description A fully funded PhD position is available at Department of Life Sciences at University of Coimbra

PhD student (f/m/d)

. They conduct research for the efficient and sustainable utilisation of crop diversity. PhD student (f/m/d) The Cryo- and Stress Biology research group is a central part of the Federal Centre Ex-situ Genebank

PhD Student (f/m/d)

supply of food, energy and raw materials, thereby addressing a major global ecological challenge. PhD Student (f/m/d) The research group Metabolic Diversity  focusses on the biochemistry and analysis

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